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Q11:
Describe transpiration pull model of water transport in plants. What are the factors influencing transpiration? How is it useful to plants?

Solution :

Step 1: The Transpiration Pull Model (Cohesion-Tension Theory)

The movement of water through the xylem is explained by the Cohesion-Tension-Transpiration Pull Model, proposed by Dixon and Joly (1894). Water movement from roots to the highest canopy is a passive process driven by the potential energy gradient created by transpiration.

  • Cohesion: Mutual attraction between water molecules due to hydrogen bonding. [Per the properties of polar molecules].
  • Adhesion: Attraction of water molecules to the polar surfaces of the xylem tracheary elements (tracheids and vessels).
  • Surface Tension: Water molecules are more attracted to each other in the liquid phase than to water in the gas phase.

As water evaporates from the mesophyll cell surfaces into the intercellular spaces and then through the stomata [Transpiration], it creates a negative pressure (tension) in the xylem sap. This tension pulls the continuous water column upward, resisting gravity.

Transpiration Xylem Conduit Water Column

Step 2: Factors Influencing Transpiration

Transpiration rates are governed by both environmental and physiological variables:

Factor Effect on Transpiration Mechanism
Temperature Directly Proportional Increases kinetic energy of water vapor; decreases relative humidity.
Light Directly Proportional Stimulates stomatal opening [via proton pumping in guard cells].
Humidity Inversely Proportional Higher ambient water vapor reduces the diffusion gradient ($\Delta C$).
Wind Speed Directly Proportional Removes the boundary layer of moist air, maintaining a steep gradient.

Step 3: Biological Significance of Transpiration

Transpiration is not merely an "unavoidable evil" but a functional necessity for plant survival:

  • Mineral Transport: It facilitates the upward movement of minerals and ions from the soil to various plant parts [Per the Mass Flow Hypothesis].
  • Thermoregulation: It exerts a cooling effect on leaf surfaces, often by as much as $10-15^\circ C$ via evaporative cooling.
  • Turgor Maintenance: By maintaining a steady flux of water, it helps keep cells turgid, which is essential for structural support and the expansion of leaves and stems.
  • Photosynthesis Supply: It ensures that the photosynthetic machinery (chloroplasts) receives a constant supply of water required for the photolysis of water ($2H_2O \rightarrow 4H^+ + 4e^- + O_2$).

Final Solution: The Transpiration Pull Model posits that negative pressure generated by water evaporation from leaves pulls a continuous, cohesive column of water through the xylem. Influenced by temperature, light, humidity, and wind, this process is essential for nutrient translocation, plant cooling, and maintenance of cell turgor pressure.


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